Literature Cited

Ardi, W., Campos-Dominguez, L., Chung, K.F., Dong, W.K., Drinkwater, E., Fuller, D., Gagul, J., Garnett, G., Girmansyah, D., Goodall-Copestake, W., Hughes, M., Jacques,E., Muaoz, O.A.J., Sang, J., Kidner, C., Kiew, R., Krishna, N., Li, R., Kumarage, L., Maw, M.B., Lin, C.W., Moonlight, P., Nguyen, H.Q., Nguyen, H.Q., Putthai, T., Pradeep, A.K., Rajbhandary, S., Rubite, R., Scherberich, D., Souvannakhoummane, K., Sreenath, M., Tebbitt, M., Thomas, D., Tian, D., Tseng, Y.H., Wilson, H. 2022 Resolving phylogenetic and taxonomic conflict in Begonia. Edinb. J. Bot 79: 1–28.
DOI: 10.24823/ejb.2022.1928View Article Google Scholar

Basumatary, S., Baruah, S., Islam, N. 2025 Begonia ovatifolia var. raimonaensis (Begoniaceae), a new variety from Northeast India. Natl. Acad. Sci. Lett. 2025: https://doi.org/10.1007/s40009-025-01663-w
DOI: 10.1007/s40009-025-01663-wView Article Google Scholar

Bora, D., Borah, D., Hughes, M., Das, S., Banik, B., Debbarma, S. Datta, B.K. 2024 Begonia tripurensis (Begoniaceae, sect. Platycentrum), a new species from Tripura, northeast India. Phytotaxa 640(2): 177–182.
DOI: 10.11646/phytotaxa.640.2.6View Article Google Scholar

Borah, D., Chowlu, K., Taram, M., Kabeer, A.A., Hughes, M., Shenoy, A. 2025 Begonia himalaica, a new species from the Indian Eastern Himalayas. Nord. J. Bot. 2025(8): e04745.
DOI: 10.1002/njb.04745View Article Google Scholar

Borah, D., Taram, M., Yama, L., Wahlsteen, E. 2021a Begonia arunachalensis (Begoniaceae), a new tuberous species of Begonia from Arunachal Pradesh, India. Gard. Bull. Singapore 73(2): 475–480.
DOI: 10.26492/gbs73(2).2021-17View Article Google Scholar

Borah, D., Taram, M., Wahlsteen, E. 2021b A new Begonia species from Arunachal Pradesh, and some notes on Begonia scintillans. Taiwania 66(4): 450–454.
DOI: 10.6165/tai.2021.66.450View Article Google Scholar

Borah, D., Taram, M., Wahlsteen, E. 2021c Begonia dicressine (Begoniaceae): a new record for India. Nat. Conserv. Res 6(4): 110–111.
DOI: 10.24189/ncr.2021.044View Article Google Scholar

Borah, D., Taram, M., Wahlsteen, E. 2021d Begonia limprichtii Irmsch. (Begoniaceae) – a new record for India. Biodiv. Res. Conserv. 61(2): 29–33.
DOI: 10.2478/biorc-2021-0002View Article Google Scholar

Borah, D., Taram, M., Hughes, M. 2023 Begonia egamii sp. nov. (Begoniaceae), a remarkable new species from Arunachal Pradesh, India. Candollea 78(2): 147–151.
DOI: 10.15553/c2023v782a5View Article Google Scholar

Borah, D., Taram, M., Das, N., Hughes, M. 2024a Begonia bijantiae, a new species of Begonia from Arunachal Pradesh, northeast India and notes on Begonia ovatifolia. Nord. J. Bot. 2024(12): e04493.
DOI: 10.1111/njb.04493View Article Google Scholar

Borah, D., Taram, M., Tasser, M. Hughes, M. 2024b A new species Begonia ziroensis and a new record of Begonia siamensis from Arunachal Pradesh, northeast India. New Zealand J. Bot. 63(1): 45–52.
DOI: 10.11646/phytotaxa.494.3.2View Article Google Scholar

Camfield, R., Hughes, M. 2018 A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India. Eur. J. Taxon. 396: 1–25.
DOI: 10.5852/ejt.2018.396View Article Google Scholar

Chen, W.-H., Huang, S.-H., Aung, A., Yuan, H.-K., Shui, Y.-M. 2019 Species diversity and spatial distribution of Begonia L. (Begoniaceae) in the Daweishan Range, Yunnan Province, China. Phytotaxa 415(1): 1–31.
DOI: 10.11646/phytotaxa.415.1.1View Article Google Scholar

Das, B., Saikia, J., Konwar, P., Siga, A. Banik, D. 2022 A new dioecious species of Begonia (Begoniaceae) from Arunachal Pradesh, India. Phytotaxa 575(1):89–96.
DOI: 10.11646/phytotaxa.575.1.6View Article Google Scholar

Doorenbos, J., Sosef, M.S., de Wilde, J.J.F.E. 1998 The Sections of Begonia: Including descriptions, keys and species Lists (Studies in Begoniaceae VI). Agricultural University, Wageningen.

Goodall-Copestake, W.P., P?rez-Espona, S., Harris, D.J., Hollingsworth, P.M. 2010 The early evolution of the mega-diverse genus Begonia (Begoniaceae) inferred from organelle DNA phylogenies. Biol. J. Linn. Soc. 101(2): 243–250.
DOI: 10.1111/j.1095-8312.2010.01489.xView Article Google Scholar

Grierson, A.J.C. 1991 Begoniaceae. In: Grierson, A.J.C., Long, D.G. (eds.), Flora of Bhutan, Vol. 2, pp. 237–245. Royal Botanic Garden, Edinburgh.

Gu, C., Peng, C.I., Turland, N.J. 2007 Begoniaceae. In Wu, C.Y., Raven, P.H. (eds.) Flora of China, Vol. 13: pp. 153–207. Science Press, Beijing & Missouri Botanical Garden Press, St. Louis.

Halbritter, H., Ulrich, S., Gr?msson, F., Weber, M., Zetter, R., Hesse, M., Buchner, R., Svojtka, M., Frosch-Radivo, A. 2018 Illustrated Pollen Terminology. Springer.
DOI: 10.1007/978-3-319-71365-6View Article

Hughes, M., Girmansyah, D. 2011. A revision of Begonia sect. Sphenanthera (Hassk.) Warb. From Sumatra. Gard. Bull. Singapore 62(2): 27–39.

Hughes, M. Moonlight, P.W., Jara, A., Tebbit, M., Wilson H., Pullan, M. 2015–Present Begonia Resource Centre. https://padme.rbge.org.uk/Begonia/ [accessed 22 October 2025].

IUCN 2024 Guidelines for using IUCN red list categories and criteria. ver.16. Prepared by the Standards and Petitions Committee.

Krishna, N., Pradeep, A.K., Amrutha, A., Jayakrishnan, T. 2021 A new species and a new record of Begonia (sect. Platycentrum, Begoniaceae) from northeastern India. Phytotaxa 482(3): 261–267.
DOI: 10.11646/phytotaxa.482.3.4View Article Google Scholar

Moonlight, P.W., Ardi, W.H., Padilla, L.A., Chung, K.F., Fuller, D., Girmansyah, D., Hollands, R., Jara-Mu?oz, A., Kiew, R., Liu, W.-C., L, Y., Mahardika, A., Marasinghe, L.D.K. O'Connor, M., Peng, C.-I, P?rez, ?.J., Phutthai, T., Pullan, M., Rajbhandary, S., Reynel, C., Rubite, R., Sang, J., Scherberich, D., Shui, Y.-M., Tebbitt, M.C., Thomas, D.C., Wilson, H.P., Zaini, N.H., Hughes, M. 2018 Dividing and conquering the fastest-growing genus: towards a natural sectional classification of the mega-diverse genus Begonia (Begoniaceae). Taxon 67(2): 267–323.
DOI: 10.12705/672.3View Article Google Scholar

Odyuo, N., Sinha, B.K., Murugesan, M., Uddin, A. 2018 A new species of Begonia L. (Begoniaceae) from Dampa Tiger Reserve, Mizoram, India. Pleione 12(1): 99–104.
DOI: 10.26679/Pleione.30.6.2018.099-104View Article Google Scholar

POWO 2026 Royal Botanic Gardens, Kew. Available from: https://powo.science.kew.org (Accessed: 2 February 2026).

Peng, C.I, Ku, S.M. 2009 Begonia × chungii (Begoniaceae), a new natural hybrid in Taiwan. Bot. Stud. 50(2): 241–250.

Plana, V., Gascoigne, A., Forrest, L.L., Harris, D., Pennington, R.T. 2004 Pleistocene and pre-Pleistocene Begonia speciation in Africa. Mol. Phylogenet. Evol. 31(2): 449–461.
DOI: 10.1016/j.ympev.2003.08.023View Article Google Scholar

Punt, W., Hoen, P.P., Blackmore, S., Nilsson, S., Le Thomas, A. 2007 Glossary of pollen and spore terminology. Rev. Palaeobot. Palynol. 143(1-2): 1–81.
DOI: 10.1016/j.revpalbo.2006.06.008View Article Google Scholar

Rajbhandary, S., Hughes, M. Shrestha, K.K. 2012 Pollen morphology of Begonia L. (Begoniaceae) in Nepal. Bangladesh J. Plant Taxon. 19(2): 191–200.
DOI: 10.3329/bjpt.v19i2.13134View Article Google Scholar

Shui, Y.M., Peng, C.I., Wu, C.Y. 2002 Synopsis of the Chinese species of Begonia (Begoniaceae), with a reappraisal of sectional delimitation. Bot. Bull. Acad. Sin. 43(4): 313–327.
DOI: 10.7016/BBAS.200210.0313View Article Google Scholar

Singh, N.P., Singh, K.P., Singh, D.K. 2002 Flora of Mizoram Vol. 1. Botanical Survey of India, Ministry of Environment and Forests, Kolkata.

Soni, A.K., Shenoy, A., Kumar, A., Anu, M., Borah, D. Kabeer, A.A. 2025 Begonia nyishiorum (Begoniaceae), a new species from Arunachal Pradesh, northeast India. Novon 33: 83–88.
DOI: 10.3417/2025991View Article Google Scholar

Taram, M., Borah, D., Hughes, M. 2021 Begonia kekarmonyingensis (Begoniaceae), a new species from Arunachal Pradesh, northeast India. Phytotaxa 494(3): 268–272.
DOI: 10.11646/phytotaxa.494.3.2View Article Google Scholar

Taram, M., Borah, D., Krishna, N., Pradeep, A.K., Amrutha, A., Hughes, M. 2020 Begonia oyuniae (Begonia sect. Monophyllon, Begoniaceae), a remarkable new species from Northeast India. Gard. Bull. Singapore 72(1): 109–115.
DOI: 10.26492/gbs72(1).2020-10View Article Google Scholar

Tebbitt, M.C. 2003 Taxonomy of Begonia longifolia and related species. Brittonia 55(1): 19–29.
DOI: 10.1663/0007-196X(2003)055[0019:TOBLBB]2.0.CO;2View Article Google Scholar

Tebbitt, M.C. 2005 Begonias: Cultivation, Identification, and Natural History. Timber Press, Portland.

Thomas, D.C., Hughes, M., Phutthai, T., Rajbhandary, S., Rubite, R., Ardi, W.H. Richardson, J.E. 2011 A non-coding plastid DNA phylogeny of Asian Begonia (Begoniaceae): evidence for morphological homoplasy and sectional polyphyly. Mol. Phylogenet. Evol. 60(3): 428–444.
DOI: 10.1016/j.ympev.2011.05.006View Article Google Scholar

Tian, D.K., Wang, W.G., Dong, L.N., Xiao, Y., Zheng, M.M., Ge, B.J. 2021 Begonia giganticaulis, a new species of Begoniaceae from southern Xizang (Tibet), China. PhytoKeys 187: 189–205.
DOI: 10.3897/phytokeys.187.75854View Article Google Scholar

Uddin, A. 2007 Distribution and status of Indian Begonia L. species. J. Econ. Taxon. Bot. 31: 591–597.

Wahlsteen, E. 2018 Begonia mariachristinae (Begoniaceae), a new species from northern Myanmar. Edinb. J. Bot. 75(2): 161–166.
DOI: 10.1017/S0960428618000021View Article Google Scholar

Wahlsteen, E. 2019 A new species of Begonia L. (Begoniaceae) and some notes on Begonia difformis from Kachin, northern Myanmar. Phytotaxa 420(3): 241–248.
DOI: 10.11646/phytotaxa.420.3.4View Article Google Scholar

Wang, W.-G., Li, R.-K., Xi, H.-C., Ma, X.-D., Shen, J.-Y. 2025 Taxonomic studies on Begonia (Begoniaceae) in Yunnan I: Two new species from southeastern Yunnan, China. Taiwania 70(3): 453?458.
DOI: 10.6165/tai.2025.70.453View Article Google Scholar